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Rhynchospora rugosa subsp. lavarum

(Gaudich.) T.Koyama (1989)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Poales Family: Cyperaceae Genus: Rhynchospora

pu‘uko‘a

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Description

Key Characters:

Growth Form: Perennials without conspicuous rhizomes.

Stems: culms tufted, 20–70 cm tall, 0.5–0.8 mm wide, 1–2–nodose below inflorescence.

Roots:

Leaves: Leaves simple. Leaves few, basal, 1–2 cauline. Blades linear, 8–30 cm long, 0.8–2.5 mm wide. Margins entire. Veins parallel. Sheaths up to 4 cm long, green, the basal ones pale brown. Stipules absent.

Flowers: Flowers in inflorescences of 2–5 small, anthelate partial panicles widely spaced on the upper ⅔ of the culm, the partial panicles solitary at a node, open, corymbiform, with 3–5 filiform branches and clusters of 2–6 subsessile spikelets, 0.8–3 cm long, bracts linear to setaceous, the lower 1–2 longer than the inflorescence, sheathing at base; spikelets brown to reddish brown, ovoid–ellipsoid, subterete, 4–5 mm long, ca. 2 mm wide; glumes 6–8, the middle 3–4 flower–bearing, 2–3 of them perfect, membranous, ovate, apex acute, the slender midnerve prolonged into a short recurved awn, the flower–bearing glumes 3–3.5 mm long, ca. 2 mm wide. hypogynous bristles shorter than the achene. Flowers all bisexual (perfect) or sometimes distalmost staminate. Perianth bristles 6, 2.5–2.7 mm long, longer than the achene (incl. the style base), antrorsely spinulose. Corolla (petals) absent. Ovary superior; styles base deltoid, 0.5–0.6 mm long, ca. 0.5 mm wide at base, gradually narrowed upward to a filiform style.

Fruit: Achenes reddish brown; obovate to obovate–elliptic; ca. 1.5 mm long; 1–1.1 mm wide; faintly transversely rugulose. Seeds 1; seed coat thin; free from pericarp; embryo basal; endosperm abundant.

Ploidy:

Habitat: Occurring in wet forest and bogs; occasionally in mesic forest.

Elevation Range: 280–1,620 m.

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Indigenous

Island Status

Kaua'i Indigenous
O'ahu Indigenous
Molokai Indigenous
Maui Indigenous
Hawai'i Indigenous

Dispersal Agents


Pollinators

Bibliography

Name Published In: Bishop Mus. Occas. Pap. 29: 127 (1989)

Other References

Wagner et al. 1990:1429 (K, O, Mo, M, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Rhynchospora rugosa subsp. lavarum 3 mi above Kaholuamano trail; with Metrosideros, Pelea, Astelia Preserved_Specimen Nagata, K.M. 2139 Kauai BISH 1980-09-09
2 Rhynchospora rugosa subsp. lavarum Site 23 OS; Piihonua; Hilo Forest Reserve Stunted `ohi`a wetland dieback. Associated spp.: Metrosideros polymorpha, Ilex anomala, Gouldia sp., Pelea clusiifolia, Cheirodendron trigynum, Myrsine lessertiana. Soil a shallow poorly drained organic found over a pahoehoe flow. Preserved_Specimen Adee, K.T. Hawaii BISH 1978-09-27
3 Rhynchospora rugosa subsp. lavarum On the trail to Site 02 OS; Piihonua; Hilo Forest Reserve Stunted `ohi`a wetland dieback. Associated spp.: Metrosideros polymorpha, Ilex anomala, Gouldia sp., Pelea clusiifolia, Cheirodendron trigynum, Myrsine lessertiana. Soil a shallow poorly drained organic found over a pahoehoe flow. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-05
4 Rhynchospora rugosa subsp. lavarum Castle Trail Preserved_Specimen Sohmer, S.H. 6068 Oahu BISH 1969-09-06
5 Rhynchospora rugosa subsp. lavarum Honaumau moist rocks Preserved_Specimen Hitchcock, A.S. 14875 Hawaii BISH 1916-04-19
6 Rhynchospora rugosa subsp. lavarum Site 23 OS; Piihonua; Hilo Forest Reserve Stunted `ohi`a wetland dieback. Assoc. spp.: Metrosideros polymorpha, Ilex anomala, Myrsine lessertiana, M. sandwicensis, Cheirodendron trigynum & Pelea clusiifolia. Soil a moderately deep but poorly drained organic/ash found on a pahoehoe flow. Preserved_Specimen Adee, K.T. Hawaii BISH 1978-09-26
7 Rhynchospora rugosa subsp. lavarum S Hilo, upper Waiakea FR, power line road, pole 27, ~2 mi S of jct. with Saddle road silversword bog; with Plantago, Carex, Lycopodium, Juncus, Sphagnum, Vaccinium Preserved_Specimen Gustafson, R. 2389 Hawaii BISH 1981-07-21
8 Rhynchospora rugosa subsp. lavarum Abundant on bog near Site 28 OS; Piihonua; Hilo Forest Reserve Primary vegetation: Metrosideros polymorpha (sometimes pubescent), Ilex anomala, Cheirodendron trigynum, Myrsine lessertiana, Vaccinium calycinum & Sadleria pallida. Soil a deep moderately well drained ash. Preserved_Specimen Adee, K.T. Hawaii BISH 1978-11-30
9 Rhynchospora rugosa subsp. lavarum South Hilo District, Waiakea Collected in wet forest off Stainback Highway. Preserved_Specimen Funk, E.J. Hawaii BISH 1998-10-19
10 Rhynchospora rugosa subsp. lavarum Wahiawa Bog Preserved_Specimen Flynn, T.W. 589 Kauai BISH 1983-09-07
11 Rhynchospora rugosa subsp. lavarum Wahiawa Bog Preserved_Specimen Wagner, W.L. 4976 Kauai BISH 1983-08-27
12 Rhynchospora rugosa subsp. lavarum Kilauea Volcano Preserved_Specimen Rock, J.F.C. 12752 Hawaii BISH 1911-04-01
13 Rhynchospora rugosa subsp. lavarum Kahakuloa, at proposed forestry planting site Preserved_Specimen Medeiros, A.C. 358 Maui BISH 1983-01-07
14 Rhynchospora rugosa subsp. lavarum [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen (Not on sheet) BISH
15 Rhynchospora rugosa subsp. lavarum Piihonua; large boggy area off Wailuku River Road Preserved_Specimen Adee, K.T. Hawaii BISH
16 Rhynchospora rugosa subsp. lavarum Oahu Preserved_Specimen Didrichsen, D.F. 3462 Oahu BISH 1845-01-01
17 Rhynchospora rugosa subsp. lavarum Konahuanui, above Manoa, lower slopes Preserved_Specimen Heller, A.A. 2343 Oahu BISH 1985-05-13
18 Rhynchospora rugosa subsp. lavarum [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen Hillebrand, W.B. BISH
19 Rhynchospora rugosa subsp. lavarum Kalalau Trail Preserved_Specimen Forbes, C.N. 81 Kauai BISH 1909-07-19
20 Rhynchospora rugosa subsp. lavarum Koolau Mts., Laie Summit Preserved_Specimen Stemmermann, R.L. 1199 Oahu BISH 1976-10-30
21 Rhynchospora rugosa subsp. lavarum Alakai Swamp Preserved_Specimen Forbes, C.N. 874 Kauai BISH 1917-07-03
22 Rhynchospora rugosa subsp. lavarum Alakai Swamp Preserved_Specimen Forbes, C.N. 874 Kauai BISH 1917-07-03
23 Rhynchospora rugosa subsp. lavarum Alakai Swamp Preserved_Specimen Forbes, C.N. 874 Kauai BISH 1917-07-03
24 Rhynchospora rugosa subsp. lavarum Kilauea Volcano Preserved_Specimen Rock, J.F.C. 8737 Hawaii BISH 1911-04-01
25 Rhynchospora rugosa subsp. lavarum Wahiawa Swamp Preserved_Specimen Forbes, C.N. 183 Kauai BISH 1909-08-01
26 Rhynchospora rugosa subsp. lavarum Nahiku Preserved_Specimen Rock, J.F.C. 8748 Maui BISH 1911-05-08
27 Rhynchospora rugosa subsp. lavarum Kilauea Volcano Preserved_Specimen Rock, J.F.C. 12752 Hawaii BISH 1911-04-01
28 Rhynchospora rugosa subsp. lavarum Hawaii Preserved_Specimen Raynal, J. 18239 Hawaii BISH 1973-08-01
29 Rhynchospora rugosa subsp. lavarum Waialua Preserved_Specimen Forbes, C.N. 401 Molokai BISH 1912-08-01
30 Rhynchospora rugosa subsp. lavarum [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen (Not on sheet) BISH
31 Rhynchospora rugosa subsp. lavarum Alakai Swamp Preserved_Specimen Forbes, C.N. 874 Kauai BISH 1917-07-03
32 Rhynchospora rugosa subsp. lavarum Kawailoa Preserved_Specimen Forbes, C.N. 2098 Oahu BISH 1915-03-02
33 Rhynchospora rugosa subsp. lavarum Koolau Mts., ~1/2 hr. Poamoho side of Castle Trail Jct Preserved_Specimen Stemmermann, R.L. 1210 Oahu BISH 1976-10-31
34 Rhynchospora rugosa subsp. lavarum West Maui, Kapunakea. Preserved_Specimen Hughes, G.D. 606 Maui BISH 1993-02-24
35 Rhynchospora rugosa subsp. lavarum Kilauea Volcano Preserved_Specimen Rock, J.F.C. 8737 Hawaii BISH 1911-04-01
36 Rhynchospora rugosa subsp. lavarum N Puu Kukui, below Haelaau Preserved_Specimen Skottsberg, C.J.F. 2789 Maui BISH 1938-08-02
37 Rhynchospora rugosa subsp. lavarum Kalalau Trail Preserved_Specimen Forbes, C.N. 81 Kauai BISH 1909-07-19
38 Rhynchospora rugosa subsp. lavarum Kahuamana swamp Preserved_Specimen Forbes, C.N. 408 Kauai BISH 1909-09-01
39 Rhynchospora rugosa subsp. lavarum Kilauea iki Preserved_Specimen Skottsberg, C.J.F. 1889 Hawaii BISH 1926-09-20
40 Rhynchospora rugosa subsp. lavarum Oahu Preserved_Specimen Mann, H. 219 Oahu BISH 1864-01-01
41 Rhynchospora rugosa subsp. lavarum Kilauea Volcano Preserved_Specimen Rock, J.F.C. 8737 Hawaii BISH 1911-04-01
42 Rhynchospora rugosa subsp. lavarum Near Honolii stream area. Preserved_Specimen Higashino, P.K. 6883 Hawaii BISH
43 Rhynchospora rugosa subsp. lavarum Pupukea Preserved_Specimen Meebold, A. Oahu BISH 1932-05-01
44 Rhynchospora rugosa subsp. lavarum North fork of Wailuku River Preserved_Specimen Adee, K.T. Hawaii BISH 1978-04-17
45 Rhynchospora rugosa subsp. lavarum Manoa, ridge toward Palolo Preserved_Specimen Meebold, A. Oahu BISH 1932-06-01
46 Rhynchospora rugosa subsp. lavarum Oahu Preserved_Specimen Gray, A. Oahu BISH
47 Rhynchospora rugosa subsp. lavarum Oahu Preserved_Specimen Andersson, N.J. Oahu BISH 1852-01-01
48 Rhynchospora rugosa subsp. lavarum [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen Hillebrand, W.B. BISH
49 Rhynchospora rugosa subsp. lavarum Aiea Hgts trail Preserved_Specimen Kerr, M. Oahu BISH 1945-12-22
50 Rhynchospora rugosa subsp. lavarum Just across Wailuku River; Area H Preserved_Specimen Adee, K.T. Hawaii BISH
51 Rhynchospora rugosa subsp. lavarum Konahuanui, above Manoa, lower slopes Preserved_Specimen Heller, A.A. 2343 Oahu BISH 1985-05-13
52 Rhynchospora rugosa subsp. lavarum Alakai Swamp Preserved_Specimen Forbes, C.N. 1702 Kauai BISH 1917-07-03
53 Rhynchospora rugosa subsp. lavarum Stainback Highway, South Hilo District Roadside in disturbed closed Metrosideros forest. Preserved_Specimen Segde. Height 2-3 ft. Clarke, G. 239 Hawaii BISH 1979-07-11
54 Rhynchospora rugosa subsp. lavarum Drive down Kona-Hilo Highway about 4 miles past Hongo Store towards Hilo. Turn right on Fern Estates Road and drive about 1.1 miles. Plant was about 25 meters east (left side) of road. (plot 56) Closed Metrosideros - Cibotium forest community Preserved_Specimen Sedge about 1.5 feet tall Newell, C.L. 654 Hawaii BISH 1966-08-28
55 Rhynchospora rugosa subsp. lavarum Hanalea District; Hanalea National Wildlife Refuge. Foothills below Kaukaopua Secondary forest of Psidium, Syzygium, and Rhodomyrtus with remnant Pandanus, Acacia, Wikstroemia, and an understory of Odontosoria, Setaria, and Lindsaea. Preserved_Specimen Clump-forming herb, extremely localized. Flynn, T.W. 6166 Kauai BISH 1997-06-06
56 Rhynchospora rugosa subsp. lavarum Alakai, between Lehua makanoe and Kilohana bog pools Preserved_Specimen Cranwell, L.M. 2952 Kauai BISH 1938-08-15
57 Rhynchospora rugosa subsp. lavarum Poamoho trail, Paalaa-Wahiawa sloping, side of trail Preserved_Specimen Wilson, K.A. 135 Oahu BISH 1952-04-06
58 Rhynchospora rugosa subsp. lavarum Located down Chain of Craters road about 9 miles toward Kalapana from intersection with Crater Rim Road; or 6.4 miles from intersection with Hilina Pali Road Metrosideros - Dicranopteris transition community Preserved_Specimen Sedge about 1 foot tall Newell, C.L. 559 Hawaii BISH 1966-08-23
59 Rhynchospora rugosa subsp. lavarum Site 30 OS; Papaikou; Hilo Forest Reserve Stunted `ohi`a wetland dieback. Assoc. spp.: Metrosideros polymorpha, Myrsine sandwicencis, Vaccinium calycinum, Styphelia tameiameiae, Paspalum sp.. Soil a thick peaty surface over a poorly drained ash; water at or near surface. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-01-12
60 Rhynchospora rugosa subsp. lavarum Waimano. Central Koolau Mountains. Growing along stream bank in gulch bottom wedged between rocks. Area dominated by Metrosideros polymorpha, Cheirodendron platyphyllum, Psidium cattleianum, Antidesma platyphyllum, Pipturus albidus, Tectaria gaudichaudii, Pritchardia martii, Pluchea caroliniana, Dicranopteris linearis, Sadleria pallida, and others. Preserved_Specimen Slender, tufted sedge with narrow leaves. Glumes not equal in size. Apex of achene with persistent thickened base of style. Style divided in two. Thomas, M.K. 594 Oahu BISH 2023-08-01
61 Rhynchospora rugosa subsp. lavarum Site 23 OS; Piihonua; Hilo Forest Reserve Stunted `ohi`a wetland dieback. Primary vegetation: Metrosideros polymorpha, Ilex anomala, Myrsine lessertiana, M. sandwicensis, Cheirodendron trigynum & Pelea clusiifolia. Soil a moderately deep poorly drained organic/ash soil on pahoehoe. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-29
62 Rhynchospora rugosa subsp. lavarum Upper Olaa Forest Resrve, Puna District Growing along hunter's trail to Kulani Cone in open canopy Metrosideros - Cibotium forest. Preserved_Specimen Sedge 3' tall. Cuddihy, L.W. 645 Hawaii BISH 1980-10-22
63 Rhynchospora rugosa subsp. lavarum Located down Chain of Craters Road about 9 miles toward Kalapana from intersection with Crater Rim Road; or 6.4 miles from intersection with Hilina Pali Road Metrosideros - native shrub - Andropogon transition community Preserved_Specimen Sedge about 2 feet tall Newell, C.L. 567 Hawaii BISH 1966-08-24
64 Rhynchospora rugosa subsp. lavarum Keaiwa (TNC preserve) Kau Forest Reserve 50-70% closed canopy Acacia koa-Metrosideros polymorpha mesic to wet forest. Occasional to common along basalt stream bed, with Deschampsia nubigena Preserved_Specimen Wood, K.R. 13084 Hawaii BISH 2009-06-10
65 Rhynchospora rugosa subsp. lavarum Koolau Mts., Laie-Malaekahana ridge stream bed Preserved_Specimen tufts St.John, H. 13061 Oahu BISH 1933-04-15
66 Rhynchospora rugosa subsp. lavarum Outer side of S rim of Kaau Crater, Koolau Mts.; along trail to crater Mesic koa forest assoc. Preserved_Specimen Herbst, D.R. 978 Oahu BISH 1968-03-10
67 Rhynchospora rugosa subsp. lavarum N of Olaa swamp on lava flow Preserved_Specimen Skottsberg, C.J.F. 425 Hawaii BISH 1922-09-08
68 Rhynchospora rugosa subsp. lavarum Ahupuaa of Waiau. Central Koolau Mountains. Growing between boulders in stream with Sporobolus spp., Pipturus albidus, Ludwigia octovalvis, Metrosideros polymorpha, Deparia petersenii, Christella parasitica, Miconia crenata, Acacia koa, Peperomia latifolia, etc.. Preserved_Specimen Clumping sedge found along edges of stream and on small islands. Thomas, M.K. 695 Oahu BISH 2024-07-17
69 Rhynchospora rugosa subsp. lavarum S Kipapa Gulch ridge Preserved_Specimen Hosaka, E.Y. 523 Oahu BISH 1932-05-15
70 Rhynchospora rugosa subsp. lavarum Waipahee trail, Kamalomaloo pasture Preserved_Specimen tufts St.John, H. 23120 Kauai BISH 1947-12-29
71 Rhynchospora rugosa subsp. lavarum Kawaihau, Mt. Wekiu, Makaleha Mts ridge crest, grasses, exposed Preserved_Specimen Nagata, K.M. 2869 Kauai BISH 1984-03-18
72 Rhynchospora rugosa subsp. lavarum Pahoehoe section of 1955 flow on the trail to Site 26 OS, Piihonua, Hilo Forest Reserve Pubescent `ohi`a dieback. Primary vegetation: Metrosideros polymorpha (pubescent), Ilex anomala, Myrsine lessertiana, Cheirodendron trigynum, Coprosma ochracea & Pelea sp.. Soil an extremely shallow well drained organic found over an a`a flow. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-27
73 Rhynchospora rugosa subsp. lavarum National Park, S rim of Makaopuhi crater open ohia Preserved_Specimen Webster, G.L. 1744 Hawaii BISH 1948-06-17
74 Rhynchospora rugosa subsp. lavarum East Maui, Hana Forest Reserve, W of Waihoi Crater bog; with Deschampsia, Carex montis-eeka, Rhacomitrium lanuginosum Preserved_Specimen Harrison, B. 592 Maui BISH 1974-01-13
75 Rhynchospora rugosa subsp. lavarum Haiku; Kahoaea; Hoaryhead Range just off road from Lihue to Lawai; near Koloa Road junction Sedge in dry open slope. Preserved_Specimen Stone, B.C. 735 Kauai BISH 1955-06-15
76 Rhynchospora rugosa subsp. lavarum Koolau Mts., above Pupukea trail grassy Preserved_Specimen Pearsall, G.A. 384 Oahu BISH 1959-05-04
77 Rhynchospora rugosa subsp. lavarum Aiea Ridge trail, summit area Open, disturbed wet grassland of Axonopus fissifolius, Sacciolepis, Paspalum conjugatum, Pterolepis; border vegetation of Metrosideros polymorpha, Broussaisia, Setaria palmifolia, Dicranopteris, Hedyotis terminalis, Freycinetia, Machaerina angustifolia, Cibotium glaucum, Cheirodendron platyphyllum. Preserved_Specimen Occasional sedge. Imada, C. 20043 Oahu BISH 2004-01-06
78 Rhynchospora rugosa subsp. lavarum Nu'uanu, Kapalama ridge trail, about 2 km past the end of the housing development Wet, invasive dominated forest, growing from the compressed soil along the trail Preserved_Specimen Uncommon along trail, clump forming sedge Faccenda, K. 2853 Oahu BISH 2022-11-20
79 Rhynchospora rugosa subsp. lavarum ridge E of Mapulehu Valley rain forest, mud in trail Preserved_Specimen St.John, H. 12894 Molokai BISH 1932-12-29
80 Rhynchospora rugosa subsp. lavarum Kamakou Preserve, wet gulch west of Pu`u Kolekole, East Moloka`i. Montane wet Metrosideros forest with native shrubs and ferns. Preserved_Specimen Small sedge with reddish brown inflorescences. Pratt, L.W. 1102 Molokai BISH 1982-05-19
81 Rhynchospora rugosa subsp. lavarum E Maui, East Kopili'ula Acacia-Metrosideros, 50% open 7 m canopy, understory of uluhe mat ferns dissected by riparian veg with ohia and Cheirodendron, understory of Ilex, Clermontia arborescens, Scaevola chamissoniana, Myrsine lessertiana, Melicope, Hedyotis, Coprosma, Perrottetia, Broussaisia, shrubs of Vaccinium, Styphelia, Cyrtandra, Rubus hawaiensis, ferns and bryophytes are common; In saturated zones along drainage and boggy sites, occasional. Preserved_Specimen Terrestrial sedge Wood, K.R. 11304 Maui BISH 2005-05-03
82 Rhynchospora rugosa subsp. lavarum Hanalei District; Alakai, just west of Waialeale Metrosideros-Rhynchospora montane bogs margined by Metrosideros-Cheirodendron mmixed wet forest with matting ferns & riparian vegetation deschampsia nubigena, Vaccinium dentatum, V. calycinum, Leptecophylla tameamea, Lycopdium venustulum, Melicope clusiifolia, broussaisia arguta, Coprosma elliptica, Kadua affinis, Dianella sandwicens Preserved_Specimen Terrestrial sedge, dominant in bogs and and bog margins Wood, K.R. 14889 Kauai BISH 2012-02-08
83 Rhynchospora rugosa subsp. lavarum Koolau Mts., above Pupukea trail weedy Preserved_Specimen Pearsall, G.A. 385 Oahu BISH 1959-05-04
84 Rhynchospora rugosa subsp. lavarum N side of Saddle Road, opposite K-7 near bulldozed area Along disturbed roadway, vegetation closed canopied Metrosideros scrub. Preserved_Specimen Wagner, W.L. 4708 Hawaii BISH 1982-11-18
85 Rhynchospora rugosa subsp. lavarum S Kipapa Gulch, Waipio, Koolau Mts ridge Preserved_Specimen Bryan Jr., E.H. Oahu BISH 1932-05-15
86 Rhynchospora rugosa subsp. lavarum Koolau Mts., 1 mi beyond head of Pupukea trail wet valley, gray clay run off Preserved_Specimen Pearsall, G.A. 387 Oahu BISH 1959-05-04
87 Rhynchospora rugosa subsp. lavarum Schofield Barracks East range, military Landing Zone named Upper 72 Open, disturbed habitat surrounded by mesicmixed native/non-native forest. Preserved_Specimen Upright herb. Beachy, J. 2013120502 Oahu BISH 2013-12-05
88 Rhynchospora rugosa subsp. lavarum Located down Chain of Craters Road about 4.1 miles toward Kalapana from intersection with Crater Rim Road. Then drive up side road with cable across entrance (Escape Road) between 1 and 3 miles. Plant at left of road Metrosideros - Sadleria - Dicranopteris forest community Preserved_Specimen Sedge about 2 feet tall. Newell, C.L. 244 Hawaii BISH 1966-08-02
89 Rhynchospora rugosa subsp. lavarum Along Opaeula and Helemano fenceline. Native Koolau summit vegetation. Preserved_Specimen Small herb. Loomis, A. 334 Oahu BISH 2013-11-01
90 Rhynchospora rugosa subsp. lavarum East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, muddy wetland W of Kaihuakala Open disturbed grassland/sedgeland dominated by Paspalum conjugatum, P. urvillei, and Sacciolepis indica, with Cyperus polystachyos, Tibouchina herbacea, Kyllinga brevifolia, Ageratina adenophora. Preserved_Specimen Occasional clumping rush. Imada, C. 200258 Maui BISH 2002-11-04
91 Rhynchospora rugosa subsp. lavarum Haiku; Kahoaea; Hoaryhead Range just off road from Lihue to Lawai; near Koloa Road junction Sedge in dry open slope. Preserved_Specimen Terrestrial reed-type epidendrum; ca. 18-24 tall. Lvs distichous, firm & leathery, olive-green sometimes red-tinged. Stone, B.C. 735 Kauai BISH 1955-06-15
92 Rhynchospora rugosa subsp. lavarum Koolau Mts., Kawailoa FR, Puu Kainapuaa with Pritchardia, Metrosideros, Antidesma, Pelea, Pittosporum Preserved_Specimen Herbst, D.R. 296 Oahu BISH 1967-01-25
93 Rhynchospora rugosa subsp. lavarum Hilo entrance to Park marshy Preserved_Specimen Fagerlund, G.O. 675 Hawaii BISH 1943-07-24
94 Rhynchospora rugosa subsp. lavarum Kaumalumalu dry aa lava, with Metrosideros Preserved_Specimen Herbst, D.R. 1748 Hawaii BISH 1970-07-09
95 Rhynchospora rugosa subsp. lavarum Located down Kona-Hilo Highway 4 miles past Hongo Store towards Hilo. Turn right onto Fern Estates Road. Plant was about 200 meters down Fern Estates Road, on right side. (plot 55)` Metrosideros - Dicranopteris - Andropogon scrub forest community Preserved_Specimen Sedge about 1.5 feet tall Newell, C.L. 672 Hawaii BISH 1966-08-31
96 Rhynchospora rugosa subsp. lavarum Puu Kahili, above fork of Wahiawa Stream muddy road Preserved_Specimen sedge Stone, B.C. 776 Kauai BISH 1955-06-15
97 Rhynchospora rugosa subsp. lavarum Manana Ridge Trail, about 1/2 of the way to the summit Along trail, mostly native area, seemingly confined to edges of trail Preserved_Specimen Clump forming, frequent along trail Faccenda, K. 2168 Oahu BISH 2021-11-24
98 Rhynchospora rugosa subsp. lavarum W Maui, Eke Crater In bog turf. Preserved_Specimen Medeiros, A.C. 323 Maui BISH 1982-11-09
99 Rhynchospora rugosa subsp. lavarum East Maui, Waihoi Bog Among Oreobolus and Rhacomitrium hummocks. Preserved_Specimen Hobdy, R.W. 2584 Maui BISH 1986-07-11
100 Rhynchospora rugosa subsp. lavarum W Maui, Eke Crater In bog turf. Preserved_Specimen Medeiros, A.C. 321 Maui BISH 1982-11-09
101 Rhynchospora rugosa subsp. lavarum Wahiawa Bog, S base of Kahili, E of Hanapepe River depressed vegetation Preserved_Specimen Fosberg, F.R. 41475 Kauai BISH 1961-03-11
102 Rhynchospora rugosa subsp. lavarum Hawaii open, wet Preserved_Specimen Lee, H.A. 155 Hawaii BISH 1926-03-01
103 Rhynchospora rugosa subsp. lavarum Waihoi Valley. Base camp area in open wet areas with other sedges and grasses. Preserved_Specimen Harrison-Gagne, B. 25 Maui BISH 1972-06-15
104 Rhynchospora rugosa subsp. lavarum 4 mi SW of Glenwood along road, rain forest Preserved_Specimen Henrickson, J.S. 4068 Hawaii BISH 1969-08-06
105 Rhynchospora rugosa subsp. lavarum Drive up Kulani Prison Road about 10.3 miles toward Kulani Prison from intersection with Kona-Hilo Highway near Hilo; up Kulani Prison Road 5.3 miles towards Kulani Prison from intersection with Volcano Road; Metrosideros - Cibotium forest community Preserved_Specimen Sedge about 2 feet tall. Newell, C.L. 699 Hawaii BISH 1966-09-01
106 Rhynchospora rugosa subsp. lavarum Kealia Forest Reserve, Makaleha Mts. Transect 2A from S of Malamalama Peak to S of Leleiwi Peak Low, open marshy wet shrubland and bogland with Metrosideros polymorpha, Antidesma platyphyllum, Dicranopteris linearis. Preserved_Specimen Perennial herb forming dense clumps in marshy area. Lorence, D.H. 7363 Kauai BISH 1993-04-16
107 Rhynchospora rugosa subsp. lavarum Upper Olaa Forest Reserve, Puna District, growing along hunter's trail to Kulani Cone in open canopy Metrosideros-Cibotium forest. Preserved_Specimen Sedge 3' tall Cuddihy, L.W. 645 Hawaii BISH 1980-10-22
108 Rhynchospora rugosa subsp. lavarum Kauluwehi Swamp Preserved_Specimen Rock, J.F.C. 5148 Kauai BISH 1911-10-01
109 Rhynchospora rugosa subsp. lavarum Waialua Preserved_Specimen Forbes, C.N. 401 Molokai BISH 1912-08-01
110 Rhynchospora rugosa subsp. lavarum Kalaupapa; Puu Alii 2, near cabin Preserved_Specimen perennial herb Wysong, M.L. 714 Molokai BISH 2005-03-24
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